Morphological Description

Branchlets terete, with longitudinal ridges, glabrous; tendrils bifurcate. Leaves with caducous stipules; petiole 1.5-4 cm, glabrous; leaf blade oval, 5-10 × 4-7.5 cm, abaxially nearly glabrous, basal veins 5, lateral veins 4 pairs, veinlets slightly raised, base cordate, margin 15-17-toothed on each side, teeth acute, apex acute or mucronulate. Panicle leaf-opposed, loose, base with well-developed branches, rachis densely pubescent. Pedicel glabrous. Berry globose, ca. 1 cm in diam. Seeds obovoid, apex rounded, surface smooth, chalazal knot elliptic, ventral holes furrowed from base to apex.