Plants polygamo-dioecious. Branchlets terete, with longitudinal ridges, densely brown villous; tendrils bifurcate or unbranched, sparsely brown pilose. Leaves simple, undivided, rarely 3-lobed; stipules brown, triangular-lanceolate, 2-4 × 1-1.5 mm, membranous, glabrous or base sparsely ciliate, margin entire, apex acuminate; petiole 0.2-3 cm, brown villous; leaf blade rhombic-ovate or elliptic, 3.5-13 × 2-7 cm, abaxially sparsely brownish pilose, adaxially sparsely pubescent only on midvein, basal veins 3, lateral veins 3-5 pairs, veinlets slightly raised abaxially, inconspicuous adaxially, base usually asymmetric, truncate or broadly so, base of lower leaves rarely subrounded, margin 6-12-toothed on each side, teeth large and sharp, rarely obtuse, apex acute. Panicle leaf-opposed, loose, 2.5-5.5 cm, basal branches undeveloped; peduncle 1-2 cm, densely brownish villous. Pedicel 1.5-2 mm, glabrous. Buds oval, 1.2-1.5 mm, apex rounded. Calyx 0.1-0.2 mm, glabrous, entire. Filaments filiform, 0.5-1 mm; anthers yellow, oval or ovate-elliptic, 0.5-0.6 mm. Ovary rounded; style short; stigma expanded. Berry globose, 6-8 mm in diam. Seeds obovoid, apex retuse, chalazal knot elliptic, ventral holes elliptic and broad, furrowed upward 1/4 from base.