Morphological Description

Plants polygamo-dioecious. Branchlets terete, with conspicuous longitudinal ridges, pubescent with simple and glandular hairs; tendrils usually bifurcate or trifurcate. Leaves simple; stipules brown, ovate-lanceolate, 7-12(-14) × 3-5 mm, membranous, glabrous, entire, apex acuminate; petiole 2-6.5 cm, with glandular and simple hairs; leaf blade oval or broadly so, 5.5-16 × 5-13.5 cm, slightly 5-lobed or undivided, abaxially tomentose with long and short hairs, adaxially pubescent, basal veins 5, lateral veins 4 or 5 pairs, veinlets conspicuously raised abaxially, slightly raised adaxially, base deeply cordate, notch acute, rarely obtuse, sometimes two sides overlapping, margin with obtuse teeth, teeth sharp. Panicle loose, leaf-opposed, 5-13 cm, basal branches well developed, sometimes with a tendril below basal branch; peduncle 1.5-3.5 cm, densely pubescent with simple and glandular hairs. Pedicel 1.6-2 mm, glabrous. Buds obovoid, 1.5-2 mm, apex rounded. Calyx cupular, ca. 2 mm, glabrous, subentire. Petals galeate. Filaments filiform, 1.4-1.8 mm; anthers yellow, elliptic, ca. 0.5 mm. Ovary conical; style short; stigma expanded. Berry globose, 7-8 mm in diam. Seeds obovoid, apex rounded and retuse, chalazal knot ovate-elliptic, ventral holes obelliptic.