Morphological Description

Branchlets terete, with longitudinal ridges; tendrils bifurcate, leaf-opposed. Leaves highly variable in shape, 3-5-foliolate, or mixed with simple and unlobed, or simple and variously lobed leaves; stipules caducous; petiole 2.5-6 cm; leaf and leaflet blades adaxially nearly glabrous, or pilose and lanate, basal veins 5, lateral veins 4-6 pairs, pubescent to sometimes slightly so or nearly glabrescent, veinlets slightly raised abaxially, inconspicuous adaxially; central leaflet rhombic-elliptic or lanceolate, 5-12 × 2.5-5 cm, base cuneate, apex acute or mucronulate; lateral leaflets ovate-elliptic or ovate-lanceolate, 3.5-9 × 3-5 cm, base asymmetric, subrounded or broadly truncate, margin with 5-20 sharp teeth on each side, apex acute or acuminate; simple leaves oval or ovate-elliptic, 5-12 × 4-8 cm, base cordate, notch obtuse, margin serrate with 21-31 teeth, apex acute. Panicle leaf-opposed, loose, 5-12 cm, basal branches well developed; peduncle 1-2.5 cm. Pedicel 1.5-2.5 mm, glabrous. Buds obovoid, 1-2.5 mm, apex rounded. Calyx glabrous, margin undulate. Filaments filiform, 0.7-1 mm. Ovary oval; style short; stigma expanded. Berry black-purple, globose, 0.8-1.3 cm in diam., glaucous. Seeds obovoid, apex retuse, chalazal knot oval, ventral holes furrowed upward 1/4 from base.